Department of
Biological Sciences

Professor, PhD
Øystein Varpe

Peer-Reviewed Articles 

Stolz C, Varpe Ø, Ims RA, Sandercock BK, Stokke BG, Fossøy F. In press.
Predator–prey interactions in the Arctic: DNA metabarcoding reveals that nestling diet of snow buntings reflects arthropod seasonality
Environmental DNA. [ doi:10.1002/edn3.439 ] [ open access ]
Kunisch EH, Graeve M, Gradinger R, Flores H, Varpe Ø, Bluhm BA. 2023.
What we do in the dark: Prevalence of omnivorous feeding activity in Arctic zooplankton during polar night
Limnology and Oceanography. 68: 1835-1851. [ doi:10.1002/lno.12389 ] [ open access ] [ pdf ]
Langbehn TJ, Aarflot JM, Freer JJ, Varpe Ø. 2023.
Visual predation risk and spatial distributions of large Arctic copepods along gradients of sea ice and bottom depth
Limnology and Oceanography. 68: 1388-1405. [ doi:10.1002/lno.12354 ] [ open access ] [ pdf ]
Larsen L-H, Cusa M, Eglund-Newby S, Berge J, Renaud PE, Varpe Ø, Geoffroy M, Falk-Petersen S. 2023.
Diets of gadoid fish in Arctic waters of Svalbard fjords during the polar night
Polar Biology. 46: 783-799. [ doi:10.1007/s00300-023-03167-z ] [ open access ] [ pdf ]
Meister N, Langbehn TJ, Varpe Ø, Jørgensen C. 2023.
Blue mussels in western Norway have vanished where in reach of crawling predators
Marine Ecology Progress Series. 721: 85-101. [ doi:10.3354/meps14416 ] [ open access ] [ pdf ]
Nowicki RC, Borgå K, Gabrielsen GW, Varpe Ø. 2023.
Energy content of krill and amphipods in the Barents Sea from summer to winter: variation across species and size
Polar Biology. 46: 139-150. [ doi:10.1007/s00300-023-03112-0 ] [ open access ] [ pdf ]
Barrio IC, Ehrich D, Soininen EM, Ravolainen VT, Bueno CG, Gilg O, Koltz AM, Speed JDM, Hik DS, Mörsdorf M, Alatalo JM, Angerbjörn A, Bêty J, Bollache L, Boulanger-Lapointe N, Brown GS, Eischeid I, Giroux MA, Hájek T, Hansen BB, Hofhuis SP, Lamarre J-F, Lang J, Latty C, Lecomte N, Macek P, McKinnon L, Myers-Smith IH, Ø. PÅ, Prevéy JS, Roth JD, Saalfeld ST, Schmidt NM, Smith P, Sokolov A, Sokolova N, Stolz C, van Bemmelen R, Varpe Ø, Woodard PF, Jónsdóttir IS. 2022.
Developing common protocols to measure tundra herbivory across spatial scales
Arctic Science. 8: 638-679. [ doi:10.1139/as-2020-0020 ] [ open access ] [ pdf ]
Barth-Jensen C, Daase M, Ormańczyk MR, Varpe Ø, Kwaśniewski S, Svensen C. 2022.
High abundances of small copepods early developmental stages and nauplii strengthen the perception of a non-dormant Arctic winter
Polar Biology. 45: 675-690. [ doi:10.1007/s00300-022-03025-4 ] [ open access ] [ pdf ]
Finne EA, Varpe Ø, Durant JM, Gabrielsen GW, Poste AE. 2022.
Nutrient fluxes from an Arctic seabird colony to the adjacent coastal marine ecosystem
Polar Biology. [ doi:10.1007/s00300-022-03024-5 ] [ open access ] [ pdf ]
Kelly C, Michelsen FA, Reite KJ, Kolding J, Varpe Ø, Berset AP, Alver MO. 2022.
Capturing big fisheries data: Integrating fishers’ knowledge in a web-based decision support tool
Frontiers in Marine Science. 9. [ doi:10.3389/fmars.2022.1051879 ] [ open access ] [ pdf ]
Keogan K, Daunt F, Wanless S, Phillips RA, Alvarez D, Anker-Nilssen T, Barrett RT, Bech C, Becker Peter H, Berglund P-A, Bouwhuis S, Burr ZM, Chastel O, Christensen-Dalsgaard S, Descamps S, Diamond T, Elliott K, Erikstad K-E, Harris M, Hentati-Sundberg J, Heubeck M, Kress SW, Langset M, Lorentsen S-H, Major HL, Mallory M, Mellor M, Miles WTS, Moe B, Mostello C, Newell M, Nisbet I, Reiertsen TK, Rock J, Shannon P, Varpe Ø, Lewis S, Phillimore AB. 2022.
Variation and correlation in the timing of breeding of North Atlantic seabirds across multiple scales
Journal of Animal Ecology. 91: 1797-1812. [ doi:10.1111/1365-2656.13758 ] [ open access ] [ pdf ]
Planque B, Aarflot JM, Buttay L, Carroll J, Fransner F, Hansen C, Husson B, Langangen Ø, Lindstrøm U, Pedersen T, Primicerio R, Sivel E, Skogen MD, Strombom E, Stige LC, Varpe Ø, Yoccoz NG. 2022.
A standard protocol for describing the evaluation of ecological models
Ecological Modelling. 471: 110059. [ doi:10.1016/j.ecolmodel.2022.110059 ] [ open access ] [ pdf ]
Skogsberg E, McGovern M, Poste A, Jonsson S, Arts MT, Varpe Ø, Borgå K. 2022.
Seasonal pollutant levels in littoral high-Arctic amphipods in relation to food sources and terrestrial run-off
Environmental Pollution. 306: 119361. [ doi:10.1016/j.envpol.2022.119361 ] [ open access ] [ pdf ]
Varpe Ø, Gabrielsen GW. 2022.
Aggregations of foraging black guillemots (Cepphus grylle) at a sea-ice edge in front of a tidewater glacier
Polar Research. 41. [ doi:10.33265/polar.v41.7141 ] [ open access ] [ pdf ]
Aune M, Raskhozheva E, Andrade H, Augustine S, Bambulyak A, Camus L, Carroll J, Dolgov AV, Hop H, Moiseev D, Renaud PE, Varpe Ø. 2021.
Distribution and ecology of polar cod (Boreogadus saida) in the eastern Barents Sea: A review of historical literature
Marine Environmental Research. 166: 105262. [ doi:10.1016/j.marenvres.2021.105262 ] [ open access ] [ pdf ]
Bandara K, Varpe Ø, Maps F, Ji R, Eiane K, Tverberg V. 2021.
Timing of Calanus finmarchicus diapause in stochastic environments
Ecological Modelling. 460: 109739. [ doi:10.1016/j.ecolmodel.2021.109739 ] [ open access ] [ pdf ]
Bandara K, Varpe Ø, Wijewardene L, Tverberg V, Eiane K. 2021.
Two hundred years of zooplankton vertical migration research
Biological Reviews. 96: 1547-1589. [ doi:10.1111/brv.12715 ] [ open access ] [ pdf ]
Ejsmond A, Forchhammer M, Varpe Ø, Jónsson JE, Jørgensen C. 2021.
Nesting synchrony and clutch size in migratory birds: Capital versus income breeding determines responses to variable spring onset
American Naturalist. 198: E122-E135. [ doi:10.1086/716064 ] [ open access ] [ pdf ] [ Online supplement ]
Geoffroy M, Langbehn T, Priou P, Varpe Ø, Johnsen G, Le Bris A, Fisher JAD, Daase M, McKee D, Cohen J, Berge J. 2021.
Pelagic organisms avoid white, blue, and red artificial light from scientific instruments
Scientific Reports. 11: 14941. [ doi:10.1038/s41598-021-94355-6 ] [ open access ] [ pdf ]
Hansen ES, Sandvik H, Erikstad KE, Yoccoz NG, Anker-Nilssen T, Bader J, Descamps S, Hodges K, Mesquita MdS, Reiertsen TK, Varpe Ø. 2021.
Centennial relationships between ocean temperature and Atlantic puffin production reveal shifting decennial trends
Global Change Biology. 27: 3753-3764. [ doi:10.1111/gcb.15665 ] [ open access ] [ pdf ]
Hobbs L, Banas NS, Cohen JH, Cottier FR, Berge J, Varpe Ø. 2021.
A marine zooplankton community vertically structured by light across diel to interannual timescales
Biology Letters. 17: 20200810. [ doi:10.1098/rsbl.2020.0810 ] [ open access ] [ pdf ]
Kunisch EH, Graeve M, Gradinger R, Haug T, Kovacs KM, Lydersen C, Varpe Ø, Bluhm BA. 2021.
Ice-algal carbon supports harp and ringed seal diets in the European Arctic: evidence from fatty acid and stable isotope markers
Marine Ecology Progress Series. 675: 181-197. [ doi:10.3354/meps13834 ] [ open access ] [ pdf ]
Meyer N, Bollache L, Galipaud M, Moreau J, Dechaume-Moncharmont F-X, Afonso E, Angerbjörn A, Bêty J, Brown G, Ehrich D, Gilg V, Giroux M-A, Hansen J, Lanctot R, Lang J, Latty C, Lecomte N, McKinnon L, Kennedy L, Reneerkens J, Saalfeld S, Sabard B, Schmidt NM, Sittler B, Smith P, Sokolov A, Sokolov V, Sokolova N, van Bemmelen R, Varpe Ø, Gilg O. 2021.
Behavioural responses of breeding arctic sandpipers to ground-surface temperature and primary productivity
Science of The Total Environment. 755: 142485. [ doi:10.1016/j.scitotenv.2020.142485 ] [ pdf ]
Samplonius JM, Atkinson A, Hassall C, Keogan K, Thackeray SJ, Assmann JJ, Burgess MD, Johansson J, Macphie KH, Pearce-Higgins JW, Simmonds EG, Varpe Ø, Weir JC, Childs DZ, Cole EF, Daunt F, Hart T, Lewis OT, Pettorelli N, Sheldon BC, Phillimore AB. 2021.
Strengthening the evidence base for temperature-mediated phenological asynchrony and its impacts
Nature Ecology & Evolution. 5: 155-164. [ doi:10.1038/s41559-020-01357-0 ] [ pdf ]
Song H, Ji R, Jin M, Li Y, Feng Z, Varpe Ø, Davis CS. 2021.
Strong and regionally distinct links between ice-retreat timing and phytoplankton production in the Arctic Ocean
Limnology and Oceanography. 66: 2498-2508. [ doi:10.1002/lno.11768 ] [ pdf ]
Barth-Jensen C, Koski M, Varpe Ø, Glad P, Wangensteen OS, Præbel K, Svensen C. 2020.
Temperature-dependent egg production and egg hatching rates of small egg-carrying and broadcast-spawning copepods Oithona similis, Microsetella norvegica and Microcalanus pusillus
Journal of Plankton Research. 42: 564-580. [ doi:10.1093/plankt/fbaa039 ] [ open access ] [ pdf ]
Claireaux M, dos Santos Schmidt TC, Olsen EM, Slotte A, Varpe Ø, Heino M, Enberg K. 2020.
Eight decades of adaptive changes in herring reproductive investment: the joint effect of environment and exploitation
ICES Journal of Marine Science. 78: 631-639. [ doi:10.1093/icesjms/fsaa123 ] [ open access ]
Kunisch EH, Bluhm BA, Daase M, Gradinger R, Hop H, Melnikov IA, Varpe Ø, Berge J. 2020.
Pelagic occurrences of the ice amphipod Apherusa glacialis throughout the Arctic
Journal of Plankton Research. 42: 73-86. [ doi:10.1093/plankt/fbz072 ] [ open access ] [ pdf ]
Piltz SH, Hjorth PG, Varpe Ø. 2020.
Empirically based minimalistic model for representing seasonal phytoplankton dynamics
Marine Ecology Progress Series. 640: 63-77. [ doi:10.3354/meps13237 ] [ pdf ]
Tarroux A, Cherel Y, Fauchald P, Kato A, Love OP, Ropert-Coudert Y, Spreen G, Varpe Ø, Weimerskirch H, Yoccoz NG, Zahn S, Descamps S. 2020.
Foraging tactics in dynamic sea-ice habitats affect individual state in a long-ranging seabird
Functional Ecology. 34: 1839-1856. [ doi:10.1111/1365-2435.13632 ] [ open access ] [ pdf ]
Bandara K, Varpe Ø, Ji R, Eiane K. 2019.
Artificial evolution of behavioral and life history strategies of high-latitude copepods in response to bottom-up and top-down selection pressures
Progress in Oceanography. 173: 134-164. [ doi:10.1016/j.pocean.2019.02.006 ] [ pdf ]
Cusa M, Berge J, Varpe Ø. 2019.
Seasonal shifts in feeding patterns: Individual and population realized specialization in a high Arctic fish
Ecology and Evolution. 9: 11112-11121. [ doi:10.1002/ece3.5615 ] [ open access ] [ pdf ]
Descamps S, Ramirez F, Benjaminsen S, Anker-Nilssen T, Barrett RT, Burr Z, Christensen-Dalsgaard S, Erikstad KE, Irons DB, Lorentsen SH, Mallory ML, Robertson GJ, Reiertsen TK, Strøm H, Varpe Ø, Lavergne S. 2019.
Diverging phenological responses of Arctic seabirds to an earlier spring
Global Change Biology. 25: 4081-4091. [ doi:10.1111/gcb.14780 ] [ open access ] [ pdf ]
Hansen BB, Lorentzen JR, Welker JM, Varpe Ø, Aanes R, Beumer LT, Pedersen ÅØ. 2019.
Reindeer turning maritime: Ice-locked tundra triggers changes in dietary niche utilization
Ecosphere. 10: e02672. [ doi:10.1002/ecs2.2672 ] [ open access ] [ pdf ]
Hitchcock DJ, Andersen T, Varpe Ø, Borgå K. 2019.
Effects of maternal reproductive investment on sex-specific pollutant accumulation in seabirds: A meta-analysis
Environmental Science & Technology. 53: 7821-7829. [ doi:10.1021/acs.est.9b01296 ] [ pdf ]
Hitchcock DJ, Andersen T, Varpe Ø, Loonen MJJE, Warner NA, Herzke D, Tombre IM, Griffin LR, Shimmings P, Borgå K. 2019.
Potential effect of migration strategy on pollutant occurrence in eggs of Arctic breeding barnacle geese (Branta leucopsis)
Environmental Science & Technology. 53: 5427-5435. [ doi:10.1021/acs.est.9b00014 ] [ pdf ]
Scherrer KJN, Kortsch S, Varpe Ø, Weyhenmeyer GA, Gulliksen B, Primicerio R. 2019.
Mechanistic model identifies increasing light availability due to sea ice reductions as cause for increasing macroalgae cover in the Arctic
Limnology and Oceanography. 64: 330-341. [ doi:10.1002/lno.11043 ] [ open access ] [ pdf ]
Bandara K, Varpe Ø, Ji R, Eiane K. 2018.
A high-resolution modeling study on diel and seasonal vertical migrations of high-latitude copepods
Ecological Modelling. 368: 357-376. [ doi:10.1016/j.ecolmodel.2017.12.010 ] [ pdf ]
Carlsson P, Vrana B, Sobotka J, Borgå K, Nizzetto PB, Varpe Ø. 2018.
New brominated flame retardants and dechlorane plus in the Arctic: Local sources and bioaccumulation potential in marine benthos
Chemosphere. 211: 1193-1202. [ doi:10.1016/j.chemosphere.2018.07.158 ] [ pdf ]
Ejsmond MJ, McNamara JM, Søreide J, Varpe Ø. 2018.
Gradients of season length and mortality risk cause shifts in body size, reserves and reproductive strategies of determinate growers
Functional Ecology. 32: 2395-2406. [ doi:10.1111/1365-2435.13191 ] [ open access ] [ pdf ]
Espinasse M, Halsband C, Varpe Ø, Gislason A, Gudmundsson K, Falk-Petersen S, Eiane K. 2018.
Interannual phenological variability in two North-East Atlantic populations of Calanus finmarchicus
Marine Biology Research. 14: 752-767. [ doi:10.1080/17451000.2018.1506135 ] [ pdf ]
Hitchcock DJ, Andersen T, Varpe Ø, Borgå K. 2018.
Improving data reporting in ecotoxicological studies
Environmental Science & Technology. 52: 8061-8062. [ doi:10.1021/acs.est.8b03324 ] [ pdf ]
Record NR, Ji RB, Maps F, Varpe Ø, Runge JA, Petrik CM, Johns D. 2018.
Copepod diapause and the biogeography of the marine lipidscape
Journal of Biogeography. 45: 2238-2251. [ doi:10.1111/jbi.13414 ] [ pdf ]
Renaud PE, Daase M, Banas NS, Gabrielsen TM, Søreide JE, Varpe Ø, Cottier F, Falk-Petersen S, Halsband C, Vogedes D, Heggland K, Berge J. 2018.
Pelagic food-webs in a changing Arctic: a trait-based perspective suggests a mode of resilience
ICES Journal of Marine Science. 75: 1871-1881. [ doi:10.1093/icesjms/fsy063 ] [ open access ] [ pdf ]
Varpe Ø, Ejsmond MJ. 2018.
Trade-offs between storage and survival affect diapause timing in capital breeders
Evolutionary Ecology. 32: 623-641. [ doi:10.1007/s10682-018-9961-4 ] [ open access ] [ pdf ]
Beumer LT, Varpe Ø, Hansen BB. 2017.
Cratering behaviour and faecal C:N ratio in relation to seasonal snowpack characteristics in a High-Arctic ungulate
Polar Research. 36: 1286121. [ doi:10.1080/17518369.2017.1286121 ] [ open access ] [ pdf ]
Espinasse M, Halsband C, Varpe Ø, Gislason A, Gudmundsson K, Falk-Petersen S, Eiane K. 2017.
The role of local and regional environmental factors for Calanus finmarchicus and C. hyperboreus abundances in the Nordic Seas
Polar Biology. 40: 2363-2380. [ doi:10.1007/s00300-017-2150-z ] [ pdf ]
Fauchald P, Tarroux A, Tveraa T, Cherel Y, Ropert-Coudert Y, Kato A, Love OP, Varpe Ø, Descamps S. 2017.
Spring phenology shapes the spatial foraging behavior of Antarctic petrels
Marine Ecology Progress Series. 568: 203-215. [ doi:10.3354/meps12082 ] [ open access ] [ pdf ]
Hitchcock DJ, Varpe Ø, Andersen T, Borgå K. 2017.
Effects of reproductive strategies on pollutant concentrations in pinnipeds: a meta-analysis
Oikos. 126: 772-781. [ doi:10.1111/oik.03955 ] [ open access ] [ pdf ]
Langbehn TJ, Varpe Ø. 2017.
Sea-ice loss boosts visual search: fish foraging and changing pelagic interactions in polar oceans
Global Change Biology. 23: 5318-5330 . [ doi:10.1111/gcb.13797 ] [ open access ] [ pdf ]
Varpe Ø. 2017.
Life history adaptations to seasonality
Integrative and Comparative Biology. 57: 943-960. [ doi:10.1093/icb/icx123 ] [ open access ] [ pdf ]
Wauchope HS, Shaw JD, Varpe Ø, Lappo EG, Boertmann D, Lanctot RB, Fuller RA. 2017.
Rapid climate-driven loss of breeding habitat for Arctic migratory birds
Global Change Biology. 23: 1085-1094. [ doi:10.1111/gcb.13404 ] [ pdf ]
Williams CM, Ragland GJ, Betini G, Buckley LB, Cheviron ZA, Donohue K, Hereford J, Humphries MM, Lisovski S, Marshall KE, Schmidt PS, Sheldon KS, Varpe Ø, Visser ME. 2017.
Understanding evolutionary impacts of seasonality: An introduction to the symposium
Integrative and Comparative Biology. 57: 921-933. [ doi:10.1093/icb/icx122 ] [ open access ] [ pdf ]
Bandara K, Varpe Ø, Søreide JE, Wallenschus J, Berge J, Eiane K. 2016.
Seasonal vertical strategies in a high-Arctic coastal zooplankton community
Marine Ecology Progress Series. 555: 49-64. [ doi:10.3354/meps11831 ] [ open access ] [ pdf ]
Burr ZM, Varpe Ø, Anker-Nilssen T, Erikstad KE, Descamps S, Barrett RT, Bech C, Christensen-Dalsgaard S, Lorentsen S-H, Moe B, Reiertsen TK, Strøm H. 2016.
Later at higher latitudes: large-scale variability in seabird breeding timing and synchronicity
Ecosphere. 7: 1-12. [ doi:10.1002/ecs2.1283 ] [ open access ] [ pdf ]
Descamps S, Tarroux A, Cherel Y, Delord K, Godø OR, Kato A, Krafft BA, Lorentsen SH, Ropert-Coudert Y, Skaret G, Varpe Ø. 2016.
At-sea distribution and prey selection of Antarctic petrels and commercial krill fisheries
Plos One. 11. [ doi:10.1371/journal.pone.0156968 ]
Descamps S, Tarroux A, Lorentsen SH, Love OP, Varpe Ø, Yoccoz NG. 2016.
Large-scale oceanographic fluctuations drive Antarctic petrel survival and reproduction
Ecography. 39: 496-505. [ doi:10.1111/ecog.01659 ] [ open access ] [ pdf ]
Jin MB, Popova EE, Zhang JL, Ji RB, Pendleton D, Varpe Ø, Yool A, Lee YJ. 2016.
Ecosystem model intercomparison of under-ice and total primary production in the Arctic Ocean
Journal of Geophysical Research-Oceans. 121: 934-948. [ doi:10.1002/2015jc011183 ] [ pdf ]
Milner JM, Varpe Ø, van der Wal R, Hansen BB. 2016.
Experimental icing affects growth, mortality, and flowering in a high Arctic dwarf shrub
Ecology and Evolution. 6: 2139-2148. [ doi:10.1002/ece3.2023 ] [ open access ] [ pdf ]
Tarroux A, Weimerskirch H, Wang SH, Bromwich DH, Cherel Y, Kato A, Ropert-Coudert Y, Varpe Ø, Yoccoz NG, Descamps S. 2016.
Flexible flight response to challenging wind conditions in a commuting Antarctic seabird: do you catch the drift?
Animal Behaviour. 113: 99-112. [ doi:10.1016/j.anbehav.2015.12.021 ] [ pdf ]
Berge J, Daase M, Renaud PE, Ambrose WG, Jr., Darnis G, Last KS, Leu E, Cohen JH, Johnsen G, Moline MA, Cottier F, Varpe Ø, Shunatova N, Balazy P, Morata N, Massabuau J-C, Falk-Petersen S, Kosobokova K, Hoppe CJM, Weslawski JM, Kuklinski P, Legezynska J, Nikishina D, Cusa M, Kedra M, Wlodarska-Kowalczuk M, Vogedes D, Camus L, Tran D, Michaud E, Gabrielsen TM, Granovitch A, Gonchar A, Krapp R, Callesen TA. 2015.
Unexpected levels of biological activity during the polar night offer new perspectives on a warming Arctic
Current Biology. 25: 2555-2561. [ doi:10.1016/j.cub.2015.08.024 ] [ pdf ]
Berge J, Renaud PE, Darnis G, Cottier F, Last K, Gabrielsen TM, Johnsen G, Seuthe L, Weslawski JM, Leu E, Moline M, Nahrgang J, Søreide JE, Varpe Ø, Lønne OJ, Daase M, Falk-Petersen S. 2015.
In the dark: A review of ecosystem processes during the Arctic polar night
Progress in Oceanography. 139: 258-271. [ doi:10.1016/j.pocean.2015.08.005 ] [ pdf ]
Cohen JH, Berge J, Moline MA, Sorensen AJ, Last K, Falk-Petersen S, Renaud PE, Leu ES, Grenvald J, Cottier F, Cronin H, Menze S, Norgren P, Varpe Ø, Daase M, Darnis G, Johnsen G. 2015.
Is ambient light during the high Arctic polar night sufficient to act as a visual cue for zooplankton?
Plos One. 10. [ doi:10.1371/journal.pone.0126247 ] [ open access ] [ pdf ]
Descamps S, Tarroux A, Varpe Ø, Yoccoz NG, Tveraa T, Lorentsen SH. 2015.
Demographic effects of extreme weather events: snow storms, breeding success, and population growth rate in a long-lived Antarctic seabird
Ecology and Evolution. 5: 314-325. [ doi:10.1002/ece3.1357 ] [ open access ] [ pdf ]
Ejsmond MJ, Varpe Ø, Czarnoleski M, Kozlowski J. 2015.
Seasonality in offspring value and trade-offs with growth explain capital breeding
American Naturalist. 186: E111-E125. [ doi:10.1086/683119 ] [ pdf ]
Grigor JJ, Marais AE, Falk-Petersen S, Varpe Ø. 2015.
Polar night ecology of a pelagic predator, the chaetognath Parasagitta elegans
Polar Biology. 38: 87-98. [ doi:10.1007/s00300-014-1577-8 ] [ pdf ]
Varpe Ø, Daase M, Kristiansen T. 2015.
A fish-eye view on the new Arctic lightscape
ICES Journal of Marine Science. 72: 2532-2538. [ doi:10.1093/icesjms/fsv129 ] [ open access ] [ pdf ]
Webster CN, Varpe Ø, Falk-Petersen S, Berge J, Stübner E, Brierley A. 2015.
Moonlit swimming: vertical distributions of macrozooplankton and nekton during the polar night
Polar Biology. 38: 75-85. [ doi:10.1007/s00300-013-1422-5 ] [ pdf ]
Berge J, Cottier F, Varpe Ø, Renaud PE, Falk-Petersen S, Kwasniewski S, Griffiths C, Søreide JE, Johnsen G, Aubert A, Bjærke O, Hovinen J, Jung-Madsen S, Tveit M, Majaneva S. 2014.
Arctic complexity: a case study on diel vertical migration of zooplankton
Journal of Plankton Research. 36: 1279-1297. [ doi:10.1093/plankt/fbu059 ] [ open access ] [ pdf ]
Daase M, Varpe Ø, Falk-Petersen S. 2014.
Non-consumptive mortality in copepods: occurrence of Calanus spp. carcasses in the Arctic Ocean during winter
Journal of Plankton Research. 36: 129-144. [ doi:10.1093/plankt/fbt079 ] [ open access ] [ pdf ]
Grigor JJ, Søreide JE, Varpe Ø. 2014.
Seasonal ecology and life- history strategy of the high-latitude predatory zooplankter Parasagitta elegans
Marine Ecology Progress Series. 499: 77-88. [ doi:10.3354/meps10676 ] [ open access ] [ pdf ]
Hansen BB, Isaksen K, Benestad RE, Kohler J, Pedersen ÅØ, Loe LE, Coulson SJ, Larsen JO, Varpe Ø. 2014.
Warmer and wetter winters: characteristics and implications of an extreme weather event in the High Arctic
Environmental Research Letters. 9. [ doi:10.1088/1748-9326/9/11/114021 ] [ open access ] [ pdf ]
Jönsson M, Varpe Ø, Kozlowski T, Berge J, Kroger RHH. 2014.
Differences in lens optical plasticity in two gadoid fishes meeting in the Arctic
Journal of Comparative Physiology a-Neuroethology Sensory Neural and Behavioral Physiology. 200: 949-957. [ doi:10.1007/s00359-014-0941-z ] [ pdf ]
Lydersen C, Assmy P, Falk-Petersen S, Kohler J, Kovacs KM, Reigstad M, Steen H, Strøm H, Sundfjord A, Varpe Ø, Walczowski W, Weslawski JM, Zajaczkowski M. 2014.
The importance of tidewater glaciers for marine mammals and seabirds in Svalbard, Norway
Journal of Marine Systems. 129: 452-471. [ doi:10.1016/j.jmarsys.2013.09.006 ] [ pdf ]
Nahrgang J, Varpe Ø, Korshunova E, Murzina S, Hallanger IG, Vieweg I, Berge J. 2014.
Gender specific reproductive strategies of an Arctic key species (Boreogadus saida) and implications of climate change
PLoS ONE. 9: e98452. [ doi:10.1371/journal.pone.0098452 ] [ open access ] [ pdf ]
Sainmont J, Andersen KH, Varpe Ø, Visser AW. 2014.
Capital versus income breeding in a seasonal environment
American Naturalist. 184: 466-476. [ doi:10.1086/677926 ] [ pdf ]
Sainmont J, Gislason A, Heuschele J, Webster C, Sylvander P, Wang M, Varpe Ø. 2014.
Inter- and intra-specific diurnal habitat selection of zooplankton during the spring bloom observed by Video Plankton Recorder
Marine Biology. 161: 1931-1941. [ doi:10.1007/s00227-014-2475-x ] [ pdf ]
Daase M, Falk-Petersen S, Varpe Ø, Darnis G, Søreide JE, Wold A, Leu E, Berge J, Philippe B, Fortier L. 2013.
Timing of reproductive events in the marine copepod Calanus glacialis: a pan-Arctic perspective
Canadian Journal of Fisheries and Aquatic Sciences. 70: 871-884. [ doi:10.1139/cjfas-2012-0401 ] [ pdf ]
Ji R, Jin M, Varpe Ø. 2013.
Sea ice phenology and timing of primary production pulses in the Arctic Ocean
Global Change Biology. 19: 734-741. [ doi:10.1111/gcb.12074 ] [ pdf ]
Kraft A, Berge J, Varpe Ø, Falk-Petersen S. 2013.
Feeding in Arctic darkness: Mid-winter diet of the pelagic amphipods Themisto abyssorum and T. libellula
Marine Biology. 160: 241-248. [ doi:10.1007/s00227-012-2065-8 ] [ pdf ]
Berge J, Varpe Ø, Moline MA, Wold A, Renaud PE, Daase M, Falk-Petersen S. 2012.
Retention of ice-associated amphipods: possible consequences for an ice-free Arctic Ocean
Biology Letters. 8: 1012-1015. [ doi:10.1098/rsbl.2012.0517 ] [ open access ] [ pdf ]
Enberg K, Jørgensen C, Dunlop ES, Varpe Ø, Boukal DS, Baulier L, Eliassen S, Heino M. 2012.
Fishing-induced evolution of growth: concepts, mechanisms, and the empirical evidence
Marine Ecology. 33: 1-25. [ doi:10.1111/j.1439-0485.2011.00460.x ] [ open access ] [ pdf ]
Gabrielsen TM, Merkel B, Søreide JE, Johansson-Karlsson E, Bailey A, Vogedes D, Nygård H, Varpe Ø, Berge J. 2012.
Potential misidentifications of two climate indicator species of the marine arctic ecosystem: Calanus glacialis and C. finmarchicus
Polar Biology. 35: 1621-1628. [ doi:10.1007/s00300-012-1202-7 ] [ open access ] [ pdf ]
Renaud PE, Berge J, Varpe Ø, Lønne OJ, Nahrgang J, Ottesen C, Hallanger I. 2012.
Is the poleward expansion by Atlantic cod and haddock threatening native polar cod, Boreogadus saida?
Polar Biology. 35: 401-412. [ doi:10.1007/s00300-011-1085-z ] [ pdf ]
Varpe Ø. 2012.
Fitness and phenology: annual routines and zooplankton adaptations to seasonal cycles
Journal of Plankton Research. 34: 267-276 . [ doi:10.1093/plankt/fbr108 ] [ pdf ]
Welcker J, Beiersdorf A, Varpe Ø, Steen H. 2012.
Mass fluctuations suggest different functions of bimodal foraging trips in a central-place forager
Behavioral Ecology. 23: 1372-1378. [ doi:10.1093/beheco/ars131 ] [ open access ] [ pdf ]
Nygård H, Wallenschus J, Camus L, Varpe Ø, Berge J. 2010.
Annual routines and life history of the amphipod Onisimus litoralis: seasonal growth, body composition and energy budget
Marine Ecology Progress Series. 417: 115-126. [ doi:10.3354/meps08798 ] [ open access ] [ pdf ]
Varpe Ø. 2010.
Stealing bivalves from common eiders: Kleptoparasitism by glaucous gulls in spring
Polar Biology. 33: 359-365. [ doi:10.1007/s00300-009-0712-4 ] [ open access ] [ pdf ]
Varpe Ø, Fiksen Ø. 2010.
Seasonal plankton-fish interactions: light regime, prey phenology, and herring foraging
Ecology. 91: 311-318. [ doi:10.1890/08-1817.1 ] [ pdf ]
Vogedes D, Varpe Ø, Søreide JE, Graeve M, Berge J, Falk-Petersen S. 2010.
Lipid sac area as a proxy for individual lipid content of Arctic calanoid copepods
Journal of Plankton Research. 32: 1471-1477. [ doi:10.1093/plankt/fbq068 ] [ open access ] [ pdf ]
Berge J, Cottier F, Last KS, Varpe Ø, Leu E, Søreide J, Eiane K, Falk-Petersen S, Willis K, Nygård H, Vogedes D, Griffiths C, Johnsen G, Lorentzen D, Brierley AS. 2009.
Diel vertical migration of Arctic zooplankton during the polar night
Biology Letters. 5: 69-72. [ doi:10.1098/rsbl.2008.0484 ] [ open access ] [ pdf ]
Berge J, Renaud PE, Eiane K, Gulliksen B, Cottier FR, Varpe Ø, Brattegard T. 2009.
Changes in the decapod fauna of an Arctic fjord during the last 100 years (1908-2007)
Polar Biology. 32: 953-961. [ doi:10.1007/s00300-009-0594-5 ] [ open access ] [ pdf ]
Hanssen SA, Bustnes JO, Tveraa T, Hasselquist D, Varpe Ø, Henden JA. 2009.
Individual quality and reproductive effort mirrored in white wing plumage in both sexes of south polar skuas
Behavioral Ecology. 20: 961-966. [ doi:10.1093/beheco/arp084 ] [ open access ] [ pdf ]
Varpe Ø, Jørgensen C, Tarling GA, Fiksen Ø. 2009.
The adaptive value of energy storage and capital breeding in seasonal environments
Oikos. 118: 363-370. [ doi:10.1111/j.1600-0706.2008.17036.x ] [ pdf ]
Heino M, Baulier L, Boukal DS, Dunlop ES, Eliassen S, Enberg K, Jørgensen C, Varpe Ø. 2008.
Evolution of growth in Gulf of St Lawrence cod?
Proceedings of the Royal Society B. 275: 1111-1112. [ doi:10.1098/rspb.2007.1429 ] [ open access ] [ pdf ]
Bustnes JO, Tveraa T, Varpe Ø, Henden JA, Skaare JU. 2007.
Reproductive performance and organochlorine pollutants in an Antarctic marine top predator: the south polar skua
Environment International. 33: 911-918. [ doi:10.1016/j.envint.2007.04.010 ] [ pdf ]
Titelman J, Varpe Ø, Eliassen S, Fiksen Ø. 2007.
Copepod mating: chance or choice?
Journal of Plankton Research. 29: 1023-1030. [ doi:10.1093/plankt/fbm076 ] [ open access ] [ pdf ]
Varpe Ø, Jørgensen C, Tarling GA, Fiksen Ø. 2007.
Early is better: seasonal egg fitness and timing of reproduction in a zooplankton life-history model
Oikos. 116: 1331-1342. [ doi:10.1111/j.2007.0030-1299.15893.x ] [ open access ] [ pdf ]
Bustnes JO, Tveraa T, Henden JA, Varpe Ø, Janssen K, Skaare JU. 2006.
Organochlorines in Antarctic and Arctic avian top predators: A comparison between the south polar skua and two species of northern hemisphere gulls
Environmental Science & Technology. 40: 2826-2831. [ doi:10.1021/es051920q ] [ pdf ]
Varpe Ø, Fiksen Ø, Slotte A. 2005.
Meta-ecosystems and biological energy transport from ocean to coast: the ecological importance of herring migration
Oecologia. 146: 443-451. [ doi:10.1007/s00442-005-0219-9 ] [ pdf ]
Varpe Ø, Tveraa T. 2005.
Chick survival in relation to nest site: is the Antarctic petrel hiding from its predator?
Polar Biology. 28: 388-394. [ pdf ]
Fiksen Ø, Varpe Ø, Kaartvedt S. 2004.
Reply to Horizons Article 'Some ideas about the role of lipids in the life cycle of Calanus finmarchicus' Irigoien (2004): II
Journal of Plankton Research. 26: 981-981. [ pdf ]
Varpe Ø, Tveraa T, Folstad I. 2004.
State-dependent parental care in the Antarctic petrel: responses to manipulated chick age during early chick rearing
Oikos. 106: 479-488. [ doi:10.1111/j.0030-1299.2004.13212.x ] [ pdf ]

Book Chapters 

Varpe Ø, Bauer S. 2022.
Seasonal animal migrations and the Arctic: Ecology, diversity, and spread of infectious agents
Pages 47-76 in Arctic one health - Challenges for northern animals and people, edited by Tryland M. Springer International Publishing.
Winder M, Varpe Ø. 2020.
Interactions in plankton food webs: Seasonal succession and phenology of Baltic Sea zooplankton
Pages 162-191 in Zooplankton Ecology, edited by Teodósio MA, Barbosa AB. CRC Press.
Varpe Ø, Ejsmond MJ. 2018.
Semelparity and iteroparity
Pages 97-124 in The Natural History of Crustacea. Vol 5: Life Histories, edited by Wellborn GA, Thiel. M.. Oxford University Press.

Popular Science Writing 

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